人类学学报 ›› 2021, Vol. 40 ›› Issue (03): 469-489.doi: 10.16359/j.1000-3193/AAS.2021.0026cstr: 32091.14.j.1000-3193/AAS.2021.0026
同号文1,2,3(), 张贝1,2,3, 陈曦4, 王晓敏5, 孙吉嘉1,2,3
收稿日期:
2020-10-27
修回日期:
2020-12-22
出版日期:
2021-06-15
发布日期:
2021-06-24
作者简介:
同号文,男,研究员,第四纪哺乳动物学专业。E-mail: 基金资助:
TONG Haowen1,2,3(), ZHANG Bei1,2,3, CHEN Xi4, WANG Xiaomin5, SUN Jijia1,2,3
Received:
2020-10-27
Revised:
2020-12-22
Online:
2021-06-15
Published:
2021-06-24
摘要:
狭义(或经典)泥河湾动物群主要产自桑干河北岸的下沙沟村周边,化石来源于不同地点;该动物群被作为我国北方早更新世的代表,但其绝对年代一直饱受争议,最新古地磁测年数据是2.2-1.7 MaBP。由于受河流及断层切割影响,泥河湾盆地内部不同区域尤其是桑干河两岸的地层对比,一直是个科学难题,其根本原因在于长期以来在桑干河南岸未发现重要化石点。最近,在桑干河南岸的岑家湾台地发现了山神庙咀化石点,目前已在此地点做过9次正规发掘,出土哺乳动物化石1526件,鉴定出25个种(含未定种),其中23种属于狭义泥河湾动物群的成员。由此看出,山神庙咀动物群的时代应当与狭义泥河湾动物群的时代接近,其地质年代应当不晚于1.7 MaBP。山神庙咀遗址与小长梁遗址隔沟相望,但其化石层稍高于小长梁文化层;山神庙咀动物群的研究结果还表明,小长梁遗址年代要比先前所测得的1.36 MaBP要早;小长梁周边的史前遗址的地质年代应当再做深入研究。小长梁地区的泥河湾层底部普遍存在砾石层,但其出现的水平位置以及砾石在分选和磨圆程度等方面都各不相同。调查清楚不同剖面中底砾层的分布状况及其性质,将会对本地区地层对比研究以及推断当时泥河湾盆地的古环境都将具有十分重要意义。
中图分类号:
同号文, 张贝, 陈曦, 王晓敏, 孙吉嘉. 泥河湾盆地早更新世山神庙咀遗址动物群及其时代意义[J]. 人类学学报, 2021, 40(03): 469-489.
TONG Haowen, ZHANG Bei, CHEN Xi, WANG Xiaomin, SUN Jijia. Chronological significance of the mammalian fauna from the Early Pleistocene Shanshenmiaozui site in Nihewan Basin, northern China[J]. Acta Anthropologica Sinica, 2021, 40(03): 469-489.
图2 山神庙咀化石点与周边遗址的位置关系 1.侏罗系顶面的夷平面,其上为更新统泥河湾层;2.山神庙咀旧石器遗址(1994年发掘区域);3.山神庙咀化石点;4.小长梁遗址A区;红色箭头指示滑坡处
Fig.2 Location of SSMZ and other prehistoric site in the vicinity
图3 山神庙咀化石点实测地层剖面图(A)及逐年发掘区域图(B)(A引自同号文等[2])
Fig.3 Profile of the strata at Shanshenmiaozui (A) and the grid system of the excavated area (B) (A is after Tong et al.[2])
图4 山神庙咀旧石器点与化石点的位置高差及探沟中出露的泥河湾层底界 A.山神庙咀旧石器遗址与化石点的位置关系;B.山神庙咀化石点发掘区域及探沟位置;C.探沟中暴露的地层序列;D.探沟底部暴露的底砾层;E.探沟中暴露的泥河湾层底部
Fig.4 Relative positions of the paleolithic site and fossil location at SSMZ as well as the exposed basal gravel in the trench
图5 小长梁地区各遗址底砾层出现位置(A-C)及特征(A’-C’) A’.小长梁遗址A区及底砾层;B-B’.山神庙咀化石点及底砾层;C-C’.照坡遗址及“石铺地面”(底砾层)
Fig.5 The feature of the basal gavels (A’-C’) and their horizons (A-C) at different sites in the XCL area
图7 山神庙咀化石点发现的小哺乳动物化石 1. Allactaga sibirica,右蹠骨(Mt II+III+IV) (V 26907);2-5. Ochotonoides complicidens,2. 右前颌骨带I1-2 (V 26905.1),3.右上颌骨带P2-M2 (V 26905.2),4.右下颌骨带p3-m3 (V 26905.3),5.左下颌骨带i1和p3-m3 (V 26905.4);6. Ochotona youngi,左下颌骨带i1和p3-m3 (V 26906)
Fig. 7 Micromammal fossils from SSMZ
图8 山神庙咀大型食草动物化石及与下沙沟有关化石比较 1a-1d.草原猛犸象,桡-尺骨(V 18010.89);2a-2c.巨副驼右侧蹠骨(V 26909.1);3a-3b.巨副驼左下颌骨(V 26909.2);4.秀丽日本鹿左角基部 (V 26910),前视;5a-5b.双叉麋鹿右侧掌骨(V 26911.2);6a-6b.双叉麋鹿右m3 (V 26911.1);7.秀丽黑鹿右角(A型);8.秀丽黑鹿右角(B型);9.秀丽黑鹿右角(C型);10.双叉麋鹿左下颌骨带p2-m3 (THP16. 875);11.秀丽黑鹿左下颌骨带p2-m3 (NIH 132);12.布氏真枝角鹿右下颌骨带p2-m3 (NIH 134)(水平翻转);13.布氏真枝角鹿左下颌骨带p2-m3 (V 25966.16)。7-8和10-11为第一作者在天津自然博物馆拍摄;12为第一作者在法国自然历史博物馆拍摄;9修改自Teilhard de Chardin and Piveteau.1930: Pl VIII-11[1]。1,2,4,5,6和13产自山神庙咀;3产自石沟;其它均产自下沙沟
Fig.8 Selected large herbivore fossils from SSMZ, compared with related fossils from Xiashagou (XSG)
狭义泥河湾动物群分类沿革Classic Nihewan Fauna | 山神庙咀动物群 SSMZ Fauna | 德马尼西动物群 Dmanisi Fauna | |||||
---|---|---|---|---|---|---|---|
Teilhard de Chardin[ | Barbour et al.[ | Teilhard de Chardin and Piveteau[ | 本文1)This paper | 本文This paper | Hemmer, et al.[ | ||
Erinaceus sp. | Erinaceus cf. dealbatus | Erinaceus cf. dealbatus | - | Beremendia fissidens | |||
Siphneus sp. | Siphneus tingi | Yangia tingi | - | - | |||
Arvicola sp. | Arvicolidae gen indet. | Borsodia chinensis | - | Mimomys tornensis | |||
Alactaga cf. annulata | Allactaga sibirica | Allactaga sibirica | |||||
Hystrix sp. | - | Hystrix sp. | |||||
Lagomys plicidens | Ochotona complicidens | Ochotonoides complicidens | Ochotonoides complicidens | - | |||
Ochotona youngi | Ochotona ex gr. Ochotona lagreli minor | ||||||
Canis megamastoides | Vulpes schlosseri | Canis (Nyctereutes) sinensis | Nyctereutes sinensis | Nyctereutes sp. | - | ||
Vulpes sp. | Vulpes sp. | Vulpes chikushanensis | - | Vulpes alopecoides | |||
Canis cf. lupus | Canis sp. | Canis chihliensis | Canis chihliensis | Canis chihliensis | Canis etruscus | ||
Canis chihliensis var. palmidens | Canis palmidens | - | - | ||||
Canis chihliensis var. minor | Eucyon minor | - | - | ||||
Ursus sp. | Ursus cf. etruscus | Ursus etruscus orientalis | - | Ursus etruscus | |||
Lutra sp. | Lutra licenti | Lutra licenti | - | - | |||
Mustela pachygnatha | Eirictis pachygnatha | - | - | ||||
Meles taxus | Meles taxus | Meles cf. leucurus | Meles chiai | - | Meles sp. | ||
Machairodus sp. | Machairodus sp. | Machairodus nihowanensis | Megantereon nihowanensis | ?Megantereon nihowanensis | Megantereon cultridens | ||
Homotherium crenatidens | - | Homotherium crenatidens | |||||
Cynailurus pleistocaenicus | Cynailurus pleistocaenicus | Acinonyx sp. | Acinonyx sp. | Acinonyx pardinensis | |||
Felis sp. | Felis sp. | Felis sp. | - | - | |||
Felis (Lynx) sp. | Lynx shansius | Lynx sp. | Lynx issiodorensis | ||||
?Panthera sp. | Panthera onca georgica | ||||||
Hyaena sp. | Chasmaporthetes progressus | - | - | ||||
Hyaena ultima | Hyaena ultima | Hyaena sinensis | Pachycrocuta licenti | Pachycrocuta licenti | Pachycrocuta perrieri Pachycrocuta sp. | ||
Crocuta honanensis | - | - | |||||
Elephas cf. trogontherii | Elephas namadicus | Mammuthus trogontherii | Mammuthus trogontherii | Mammuthus meridionalis | |||
Chalicotherid gen. indet. | Chalicotheridae gen. indet. | Postschizotherium chardini | - | - | |||
Hipparion sp. | Hipparion sinense | Hipparion (Proboscidipparion) sinense | Hipparion sp. | ||||
Equus sp. | Equus sp. | Equus sanmeniensis | Equus sanmeniensis | Equus sanmeniensis | Equus stenonis | ||
Equus teilhardi | - | - | |||||
Rhinoceros sp. | Rhinoceros cf. sinensis | Stephanorhinus yunchuchenensis | - | Stephanorhinus etruscus | |||
Rhinoceros cf. tichorhinus | Rhinoceros cf. tichorhinus | Coelodonta nihowansis | Coelodonta nihowansis | - | |||
Rhinocerid gen. indet. | Elasmotherium sp. | Elasmotherium sp. | Elasmotherium peii | - | |||
Circotherium sp. | Hesperotherium sp. | - | - | ||||
Sus sp. | Sus cf. lydekkeri | Sus lydekkeri | Sus lydekkeri | - | |||
Camelus (Paracamelus) gigas | Camelus (Paracamelus) gigas | Paracamelus gigas | Paracamelus gigas | - | |||
Cervus sp. | Cervulus cf. munjack | Cervulus cf. sinensis | Muntiacus bohlini | - | - | ||
Cervus cf. rusa | Cervus (Rusa) elegans | Nipponicervus elegans | Nipponicervus elegans | Dama (=Pseudodama =Cervus s.l.) cf. nestii | |||
Cervus cf. tetraceros | Cervus (Eucladoceros) boulei | Eucladoceros boulei | Eucladoceros boulei | Eucladoceros aff. tegulensis | |||
Cervus sp. | Cervus sp. | Cervus (Elaphurus) bifurcatus | Elaphurus bifurcatus | Elaphurus bifurcatus | - | ||
Antilope gen. indet. | Spirocerus wongi | Spirocerus wongi | Spirocerus wongi | Pontoceros sp. | |||
Spirocerus cf. peii | - | ||||||
?Antilospira robusta | - | ||||||
Gazella sp. | Gazella sp. | Gazella sinensis | Gazella sinensis | Gazella sinensis | - | ||
Gazella cf. subgutturosa | Gazella subgutturosa | - | - | ||||
Antilope gen. indet. | Hydropotes sp. | - | - | ||||
Ovis sp. | Ovis shantungensis | Ovis shantungensis | Ovis shantungensis | ||||
Ovis or Capra indet. | Ovis or Capra indet. | - | Capra dalii | ||||
Euceratherium (?) | Ovibovine gen. indet. | Megalovis piveteaui | Megalovis piveteaui | Soergelia cf. minor | |||
Leptobos sp. | Bison palaeosinensis | Bison palaeosinensis | Bison palaeosinensis | Bison (Eobison) georgicus |
表1 狭义泥河湾动物群组成及与山神庙咀动物群的对比
Tab.1 The taxonomic compositions of the Classic Nihewan Fauna and SSMZ Fauna
狭义泥河湾动物群分类沿革Classic Nihewan Fauna | 山神庙咀动物群 SSMZ Fauna | 德马尼西动物群 Dmanisi Fauna | |||||
---|---|---|---|---|---|---|---|
Teilhard de Chardin[ | Barbour et al.[ | Teilhard de Chardin and Piveteau[ | 本文1)This paper | 本文This paper | Hemmer, et al.[ | ||
Erinaceus sp. | Erinaceus cf. dealbatus | Erinaceus cf. dealbatus | - | Beremendia fissidens | |||
Siphneus sp. | Siphneus tingi | Yangia tingi | - | - | |||
Arvicola sp. | Arvicolidae gen indet. | Borsodia chinensis | - | Mimomys tornensis | |||
Alactaga cf. annulata | Allactaga sibirica | Allactaga sibirica | |||||
Hystrix sp. | - | Hystrix sp. | |||||
Lagomys plicidens | Ochotona complicidens | Ochotonoides complicidens | Ochotonoides complicidens | - | |||
Ochotona youngi | Ochotona ex gr. Ochotona lagreli minor | ||||||
Canis megamastoides | Vulpes schlosseri | Canis (Nyctereutes) sinensis | Nyctereutes sinensis | Nyctereutes sp. | - | ||
Vulpes sp. | Vulpes sp. | Vulpes chikushanensis | - | Vulpes alopecoides | |||
Canis cf. lupus | Canis sp. | Canis chihliensis | Canis chihliensis | Canis chihliensis | Canis etruscus | ||
Canis chihliensis var. palmidens | Canis palmidens | - | - | ||||
Canis chihliensis var. minor | Eucyon minor | - | - | ||||
Ursus sp. | Ursus cf. etruscus | Ursus etruscus orientalis | - | Ursus etruscus | |||
Lutra sp. | Lutra licenti | Lutra licenti | - | - | |||
Mustela pachygnatha | Eirictis pachygnatha | - | - | ||||
Meles taxus | Meles taxus | Meles cf. leucurus | Meles chiai | - | Meles sp. | ||
Machairodus sp. | Machairodus sp. | Machairodus nihowanensis | Megantereon nihowanensis | ?Megantereon nihowanensis | Megantereon cultridens | ||
Homotherium crenatidens | - | Homotherium crenatidens | |||||
Cynailurus pleistocaenicus | Cynailurus pleistocaenicus | Acinonyx sp. | Acinonyx sp. | Acinonyx pardinensis | |||
Felis sp. | Felis sp. | Felis sp. | - | - | |||
Felis (Lynx) sp. | Lynx shansius | Lynx sp. | Lynx issiodorensis | ||||
?Panthera sp. | Panthera onca georgica | ||||||
Hyaena sp. | Chasmaporthetes progressus | - | - | ||||
Hyaena ultima | Hyaena ultima | Hyaena sinensis | Pachycrocuta licenti | Pachycrocuta licenti | Pachycrocuta perrieri Pachycrocuta sp. | ||
Crocuta honanensis | - | - | |||||
Elephas cf. trogontherii | Elephas namadicus | Mammuthus trogontherii | Mammuthus trogontherii | Mammuthus meridionalis | |||
Chalicotherid gen. indet. | Chalicotheridae gen. indet. | Postschizotherium chardini | - | - | |||
Hipparion sp. | Hipparion sinense | Hipparion (Proboscidipparion) sinense | Hipparion sp. | ||||
Equus sp. | Equus sp. | Equus sanmeniensis | Equus sanmeniensis | Equus sanmeniensis | Equus stenonis | ||
Equus teilhardi | - | - | |||||
Rhinoceros sp. | Rhinoceros cf. sinensis | Stephanorhinus yunchuchenensis | - | Stephanorhinus etruscus | |||
Rhinoceros cf. tichorhinus | Rhinoceros cf. tichorhinus | Coelodonta nihowansis | Coelodonta nihowansis | - | |||
Rhinocerid gen. indet. | Elasmotherium sp. | Elasmotherium sp. | Elasmotherium peii | - | |||
Circotherium sp. | Hesperotherium sp. | - | - | ||||
Sus sp. | Sus cf. lydekkeri | Sus lydekkeri | Sus lydekkeri | - | |||
Camelus (Paracamelus) gigas | Camelus (Paracamelus) gigas | Paracamelus gigas | Paracamelus gigas | - | |||
Cervus sp. | Cervulus cf. munjack | Cervulus cf. sinensis | Muntiacus bohlini | - | - | ||
Cervus cf. rusa | Cervus (Rusa) elegans | Nipponicervus elegans | Nipponicervus elegans | Dama (=Pseudodama =Cervus s.l.) cf. nestii | |||
Cervus cf. tetraceros | Cervus (Eucladoceros) boulei | Eucladoceros boulei | Eucladoceros boulei | Eucladoceros aff. tegulensis | |||
Cervus sp. | Cervus sp. | Cervus (Elaphurus) bifurcatus | Elaphurus bifurcatus | Elaphurus bifurcatus | - | ||
Antilope gen. indet. | Spirocerus wongi | Spirocerus wongi | Spirocerus wongi | Pontoceros sp. | |||
Spirocerus cf. peii | - | ||||||
?Antilospira robusta | - | ||||||
Gazella sp. | Gazella sp. | Gazella sinensis | Gazella sinensis | Gazella sinensis | - | ||
Gazella cf. subgutturosa | Gazella subgutturosa | - | - | ||||
Antilope gen. indet. | Hydropotes sp. | - | - | ||||
Ovis sp. | Ovis shantungensis | Ovis shantungensis | Ovis shantungensis | ||||
Ovis or Capra indet. | Ovis or Capra indet. | - | Capra dalii | ||||
Euceratherium (?) | Ovibovine gen. indet. | Megalovis piveteaui | Megalovis piveteaui | Soergelia cf. minor | |||
Leptobos sp. | Bison palaeosinensis | Bison palaeosinensis | Bison palaeosinensis | Bison (Eobison) georgicus |
图9 狭义泥河湾动物群组成及其地质分布 蓝色标记的属种也出现在山神庙咀化石点;橙色标记的地点有古人类化石发现;绿色区域代表狭义泥河湾动物群和山神庙咀动物群主要分布时段;各属种的分布时限主要参考邱占祥[29],Qiu[52]及Rook and Martínez-Navarro[56]
Fig.9 The up-dated taxonomic compositions of the Classic Nihewan Fauna
[1] | Teilhard de Chardin P, Piveteau J. Les mammiféres fossils de Nihowan (Chine)[M]. Annales de Paléontologie, 1930,19:1-134 |
[2] | 同号文, 胡楠, 韩非. 河北阳原泥河湾盆地山神庙咀早更新世哺乳动物群的发现[J]. 第四纪研究, 2011,31(4):643-653 |
[3] |
Tong HW. New remains of Mammuthus trogontherii from the Early Pleistocene Nihewan beds at Shanshenmiaozui, Hebei[J]. Quaternary International, 2012,255:217-230
doi: 10.1016/j.quaint.2011.07.035 URL |
[4] | Tong HW, Hu N, Wang XM. New remains of Canis chihliensis (Mammalia, Carnivora) from Shanshenmiaozui, a Lower Pleistocene Site in Yangyuan, Hebei[J]. Vertebrata PalAsiatica, 2012,50(4):335-360 |
[5] | 同号文, 王法岗, 郑敏, 等. 泥河湾盆地新发现的梅氏犀及裴氏板齿犀化石[J]. 人类学学报, 2014,33(3):369-388 |
[6] |
Tong HW, Wang XM. Juvenile skulls and other postcranial bones of Coelodonta nihowanensis from Shanshenmiaozui, Nihewan Basin, China[J]. Journal of Vertebrate Paleontology, 2014,34(3):710-724
doi: 10.1080/02724634.2013.814661 URL |
[7] |
Tong HW, Chen X. On newborn calf skulls of Early Pleistocene Mammuthus trogontherii from Shanshenmiaozui in Nihewan Basin, China[J]. Quaternary International, 2016,406:57-69
doi: 10.1016/j.quaint.2015.02.026 URL |
[8] |
Tong HW, Chen X, Zhang B. New fossils of Bison palaeosinensis (Artiodactyla, Mammalia) from the steppe mammoth site of Early Pleistocene in Nihewan Basin, China[J]. Quaternary International, 2017,445:250-268
doi: 10.1016/j.quaint.2016.07.033 URL |
[9] |
Chen X, Tong HW. On the hindfoot bones of Mammuthus trogontherii from Shanshenmiaozui in Nihewan Basin, China[J]. Quaternary International, 2017,445:50-59
doi: 10.1016/j.quaint.2016.09.001 URL |
[10] | Tong HW, Chen X, Zhang B. New postcranial bones of Elasmotherium peii from Shanshenmiaozui in Nihewan Basin, Northern China[J]. Quaternaire, 2018,29(3):195-204 |
[11] |
Tong HW, Zhang B. New fossils of Eucladoceros boulei (Artiodactyla, Mammalia) from Early Pleistocene Nihewan Beds, China[J]. Palaeoworld, 2019,28(3):403-424
doi: 10.1016/j.palwor.2019.05.003 URL |
[12] | Tong HW, Chen X, Zhang B, et al. Hypercarnivorous teeth and healed injuries to Canis chihliensis from Early Pleistocene Nihewan beds, China, support social hunting for ancestral wolves[J]. PeerJ, 2020, 8: e9858, https://doi.org/10.7717/peerj.9858 |
[13] |
Wang M, O’Connor JK, Zhou ZH. The first fossil crow (Corvus sp. indet.) from the Early Pleistocene Nihewan Paleolithic sites in North China[J]. Journal of Archaeological Science, 2013,40(3):1623-1628
doi: 10.1016/j.jas.2012.10.026 URL |
[14] | 卫奇, 李珺, 裴树文. 旧石器遗址与古人类文化[A].见:袁宝印,夏正楷,牛平山(主编). 泥河湾裂谷与古人类[M]. 北京: 地质出版社, 2011: 132-207 |
[15] | 陈茅南( 主编). 泥河湾层的研究[M]. 北京: 海洋出版社, 1988: 1-145 |
[16] | 陈曦. 泥河湾盆地早更新世山神庙咀化石地点埋藏学研究[D]. 北京:中国科学院古脊椎动物与古人类研究所, 2018: 1-140 |
[17] | Boule M, Breuil H, Licent E, et al. Le Paléolithique de la Chine[M]. Paris: Masson et Cie Editeurs, 1928: 1-138 |
[18] | Wu WY, Flynn LJ. The Lagomorphs (Ochotonidae, Leporidae) of Yushe Basin[A]. In: Flynn LJ, Wu WY (Eds.). Late Cenozoic Yushe Basin, Shanxi Province, China: Geology and Fossil Mammals, Volume II: Small Mammal Fossils of Yushe Basin[C]. Dordrecht: Springer, 2017: 31-58 |
[19] | Erbajeva MA, Zheng SH. New data on Late Miocene - Pleistocene ochotonids (Ochotonidae, Lagomorpha) from North China[J]. Acta Zoologica Cracoviensia A, 2005,48(1):93-117 |
[20] | 蔡保全, 李强, 郑绍华. 泥河湾盆地马圈沟遗址化石哺乳动物及年代讨论[J]. 人类学学报, 2008,27(2):127-140 |
[21] |
Moore TY, Organ CL, Edwards SV, et al. Multiple phylogenetically distinct events shaped the evolution of limb skeletal morphologies associated with bipedalism in the jerboas[J]. Current Biology, 2015,25:2785-2794
doi: 10.1016/j.cub.2015.09.037 URL |
[22] | Wei GB, Taruno H, Jin CZ, et al. The earliest spec,imens of the steppe mammoth, Mammuthus trogontherii from the Early Pleistocene Nihewan Formation, North China[J]. Earth Science, 2003,57:289-298 |
[23] | Wei GB, Taruno, H, Yoshinari K, et al. Pliocene and Early Pleistocene primitive mammoths of northern China: Their revised taxonomy, biostratigraphy and evolution[J]. Journal of Geosciences, Osaka City University, 2006,49(5):59-101 |
[24] | Young CC. On the Artiodactyla from the Sinanthropus site at Choukoutien[M]. Palaeontologia Sinica, Ser. C, 1932,8(2):1-100 |
[25] | Teilhard de Chardin P, Trassaert M. Pliocene Camelidae, Giraffidae and Cervidae of south-eastern Shansi[M]. Palaeontologia Sinica, New Ser. C, 1937,1:1-68 |
[26] | 陈曦, 赵海龙, 张贝, 等. 泥河湾盆地石沟遗址B区发掘报告[J]. 第四纪研究, 2017,37(4):895-907 |
[27] | Zdansky O. Paracamelus gigas, Schlosser[J]. Palaeontologia Sinica, Ser. C, 1926,2(4):1-44 |
[28] | Bohlin B. Die Familie Giraffidae[M]. Palaeontologia Sinica, Ser. C, 1927,4(1):1-178 |
[29] | 邱占祥, 邓涛, 王伴月. 甘肃东乡龙担早更新世哺乳动物群[M]. 中国古生物志新丙种第27号, 北京:科学出版社, 2004: 1-224 |
[30] | 邱占祥. 泥河湾哺乳动物群与中国第四纪下限[J]. 第四纪研究, 2000,20(2):142-154 |
[31] |
Samejima Y, Matsuoka H. A new viewpoint on antlers reveals the evolutionary history of deer (Cervidae, Mammalia)[J]. Scientific Reports, 2020, 10(1):8910, https://doi.org/10.1038/s41598-020-64555-7
doi: 10.1038/s41598-020-64555-7 URL pmid: 32488122 |
[32] |
Barbour GB. Preliminary Observations in the Kalgan Area[J]. Bulletin of the Geological Society of China, 1924,3(2):153-168
doi: 10.1111/j.1755-6724.1924.mp3002009.x URL |
[33] |
Barbour GB. The Deposits of the Sang Kan Ho Valley[J]. Bulletin of the Geological Society of China, 1925,4(1):53-55
doi: 10.1111/j.1755-6724.1925.mp4001008.x URL |
[34] | Licent E. Onze années (1923-1933) de séjour et d’exploration dans le Bassin du Fleuve Jaune, du Pai Ho et des autres tributaries du Golfe du Pei Tcheu Ly (Tome I)[M]. Tientsin: Mission de Sienhsien, 1935: 1-296 |
[35] | Licent E. Onze années (1923-1933) de séjour et d’exploration dans le bassin du Fleuve Jaune, du Paiho et des autres tributaries du Golfe du Pei Tcheu Ly (Tome II)[M]. Tientsin: Mission de Sienhsien, 1936: 297-718 |
[36] | 刘文晖. 泥河湾盆地红崖扬水站地点貉类化石及貉属化石分类的修订[D]. 北京:中国科学院古脊椎动物与古人类研究所, 2019: 1-384 |
[37] | Farjand A. Lithostratigraphy, biostratigraphy, and geochronology of Xiashagou Strata with focus on the classic Nihewan Fauna[D]. Beijing: Institute of Vertebrate paleontology and Paleoanthropology, The University of Chinese Academy of Sciences, 2020: 1-176 |
[38] |
Barbour GB, Licent E, Teilhard de Chardin P. Geological study of the deposits of the Sangkanho Basin[J]. Bulletin of the Geological Society of China, 1927,5(3-4):263-278
doi: 10.1111/j.1755-6724.1926.mp53-4005.x URL |
[39] |
Teilhard de Chardin P. Palaeontological notes[J]. Bulletin of the Geological Society of China, 1926,5(1) : 57-59
doi: 10.1111/(ISSN)1755-6724b URL |
[40] | 陈蜜, 韩琦. 泥河湾地质遗址的发现—以桑志华、巴尔博对泥河湾研究的优先权为中心[J]. 自然科学史研究, 2016,35(3):320-340 |
[41] | 李永项, 张云翔, 孙博阳, 等. 泥河湾新发现的早更新世真马化石[J]. 中国科学(地球科学), 2015,45(10):1457-1468 |
[42] | 李凯清, 岳峰, 王旭日, 等. 泥河湾动物群化石新材料[A].见:董为,张颖奇(主编).第十七届中国古脊椎动物学学术年会论文集[C].北京:海洋出版社,(待刊) |
[43] |
Liu WH, Dong W, Zhang LM, et al. New material of Early Pleistocene Sus (Artiodactyla, Mammalia) from Yangshuizhan in Nihewan Basin, North China[J]. Quaternary International, 2017,434:32-47
doi: 10.1016/j.quaint.2015.12.104 URL |
[44] | 汤英俊. 河北蔚县早更新世哺乳动物化石及其在地层划分上的意义[J]. 古脊椎动物与古人类, 1980,18(4):314-323 |
[45] | 卫奇. 在泥河湾层中发现纳玛象头骨化石[J]. 古脊椎动物学报, 1976,4(1):53-58 |
[46] | 李凯清, 赵文俭, 岳峰, 等. 河北泥河湾盆地晚新生代的长鼻类化石[A].见:董为(主编).第十五届中国古脊椎动物学学术年会论文集[C].北京:海洋出版社, 2016: 87-96 |
[47] | 汤英俊, 计宏祥. 河北蔚县上新世–早更新世间的一个过渡动物群[J]. 古脊椎动物与古人类, 1983,21(3):245-254 |
[48] | 卫奇. 泥河湾层中的大角鹿一新种[J].古脊椎动物学报, 1983(1):87-95 |
[49] | 郑绍华, 蔡保全. 河北蔚县东窑子头大南沟剖面中的小哺乳动物化石[A].见:中国科学院古脊椎动物与古人类研究所(编).第十三届国际第四纪大会论文选[C].北京:北京科学技术出版社, 1991: 100-131 |
[50] | 郑绍华, 蔡保全, 李强. 泥河湾盆地洞沟剖面上新世/更新世小哺乳动物[J]. 古脊椎动物学报, 2006,44(4):320-331 |
[51] | Qiu ZX, Qiu ZD. Chronological sequence and subdivision of Chinese Neogene mammalian faunas[J]. Palaeogeography, Palaeoclimatology, Palaeocelogy, 1995,116(1-2):41-70 |
[52] | Qiu ZX. Quaternary environmental changes and evolution of large mammals in North China[J]. Vertebrata PalAsiatica, 2006,44(2):109-132 |
[53] | 同号文, 汤英俊, 袁宝印. 脊椎动物地层划分[A].见:袁宝印,夏正楷,牛平山(主编).泥河湾裂谷与古人类[M]. 北京: 地质出版社, 2011: 47-60 |
[54] |
Hemmer H, Kahlke RD, Vekua AK. The cheetah Acinonyx pardinensis (Croizet et Jobert, 1828) s.l. at the hominin site of Dmanisi (Georgia) - A potential prime meat supplier in Early Pleistocene ecosystems[J]. Quaternary Science Reviews, 2011,30(19-20):2703-2714
doi: 10.1016/j.quascirev.2011.05.024 URL |
[55] | Liu P, Deng C, Li S, et al. Magnetostratigraphic dating of the Xiashagou Fauna and implication for sequencing the mammalian faunas in the Nihewan Basin, North China[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 2012,315:75-85 |
[56] |
Rook L, Martínez-Navarro B. Villafranchian: The long story of a Plio-Pleistocene European large mammal biochronologic unit[J]. Quaternary International, 2010,219:134-144
doi: 10.1016/j.quaint.2010.01.007 URL |
[57] | Madurell-Malapeira J, Ros-Montoya S, Espigares MP, et al. “Villafranchian large mammals from the Iberian Peninsula: paleobiogeography, paleoecology and dispersal events”[J]. Journal of Iberian Geology, 2014,40(1):167-178 |
[58] | Per C, Adolfssen JS, 2007. Osteology and ecology of Megantereon cultridens SE311 (Mammalia; Felidae; Machairodontinae), a sabrecat from the Late Pliocene - Early Pleistocene of Senéze, France[J]. Zoological Journal of the Linnean Society(4):833-884 |
[59] |
Azzaroli A. Quaternary mammals and the “End-Villafranchian” dispersal event. A turning point in the history of Eurasia[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 1983,44:117-139
doi: 10.1016/0031-0182(83)90008-1 URL |
[60] | Sardella R, Palombo MR. The Pliocene-Pleistocene boundary: which significance for the so-called Wolf event? Evidences from Western Europe[J]. Quaternaire, 2007,18:65-71 |
[61] |
Sotnikova M, Rook L. Dispersal of the Canini (Mammalia, Canidae: Caninae) across Eurasia during the Late Miocene to Early Pleistocene[J]. Quaternary International, 2010,212(2):86-97
doi: 10.1016/j.quaint.2009.06.008 URL |
[62] |
Pueyo EL, Muñoz A, Laplana C, et al. The last appearance datum of Hipparion in Western Europe: Magnetostratigraphy along the Pliocene-Pleistocene boundary in the Villarroya Basin (Northern Spain)[J]. International Journal of Earth Sciences, 2016,105(8):2203-2220
doi: 10.1007/s00531-015-1281-0 URL |
[63] | Kahlke RD. Les communautés de grands mammifères de la partie supérieure du Pléistocène inférieur et la biochrone de l’épivillafranchien[J]. Quaternaire, 2009,20(4):415-427 |
[64] |
Bellucci L, Sardella R, Rook L. Large mammal biochronology framework in Europe at Jaramillo: The Epivillafranchian as a formal biochron[J]. Quaternary International, 2015,389:84-89
doi: 10.1016/j.quaint.2014.11.012 URL |
[65] | 谢飞, 李珺, 刘连强. 泥河湾旧石器文化[M]. 石家庄: 花山文艺出版社, 2006: 1-278 |
[66] |
Gao X, Wei Q, Shen C, et al. New light on the earliest hominid occupation in East Asia[J]. Current Anthropology, 2005,46(Suppl):S115-S120
doi: 10.1086/497666 URL |
[67] |
Gabunia L, Vekua A, Lordkipanidze D, et al. Earliest Pleistocene hominid cranial remains from Dmanisi, Republic of Georgia: taxonomy, geological setting, and age[J]. Science, 2000,288(5468):1019-1025
doi: 10.1126/science.288.5468.1019 URL |
[68] |
Ferring R, Oms O, Agusti J, et al. Earliest human occupations at Dmanisi (Georgian Caucasus) dated to 1.85-1.78 Ma.[J]. Proceedings of the National Academy of Sciences of the United States of America, 2011,108(26):10432-10436
doi: 10.1073/pnas.1106638108 pmid: 21646521 |
[69] |
Liu P, Wu Z, Deng C, et al. Magnetostratigraphic dating of the Shanshenmiaozui mammalian fauna in the Nihewan Basin, North China[J]. Quaternary International, 2016,400:202-211
doi: 10.1016/j.quaint.2014.09.024 URL |
[70] |
Zhu RX, Hofrman KA, Potts R, et al. Earliest presence of humans in northeast Asia[J]. Nature, 2001,413:413-417
pmid: 11574886 |
[71] |
Deng CL, Zhu RX, Zhang R, et al. Timing of the Nihewan formation and faunas[J]. Quaternary Research, 2008,69(1):77-90
doi: 10.1016/j.yqres.2007.10.006 URL |
[72] |
Licent E, Trassaert M. The Pliocene lacustrine series in central Shansi[J]. Bulletin of the Geological Society of China, 1935,14:211-220
doi: 10.1111/(ISSN)1755-6724b URL |
[1] | 胡海虔, 黄万波, 魏光飚, 代辉, 熊璨, 何树兴, 姜涛. 重庆武隆早更新世地层中发现巨猿化石[J]. 人类学学报, 2024, 43(05): 701-711. |
[2] | 陈育芝, 武仙竹. 长江三峡及周边地区早期人类的生存环境与生存行为[J]. 人类学学报, 2024, 43(02): 287-297. |
[3] | 赵云啸, 仝广, 涂华, 赵海龙. 河北省泥河湾盆地石沟遗址C区发掘简报[J]. 人类学学报, 2024, 43(01): 122-131. |
[4] | 刘连强, 蒲昱晓, 侯佳岐, 王法岗. 泥河湾盆地马圈沟遗址鱼咀沟1号地点2017-2018年发掘出土的石制品[J]. 人类学学报, 2024, 43(01): 40-54. |
[5] | 王晓敏, 刘连强, 陈国鹏, 李锋, 谢飞, 高星. 泥河湾盆地马圈沟遗址哺乳动物破碎长骨反映的古人类行为[J]. 人类学学报, 2024, 43(01): 91-105. |
[6] | 张月书, 李锋, 陈福友, 仪明洁, 高星. 泥河湾盆地东谷坨遗址6A2层的形成过程[J]. 人类学学报, 2023, 42(01): 61-74. |
[7] | 王元, 王一飒, 王奕迪, 秦超, 秦大公, 金昌柱. 广西扶绥岩亮洞与巨猿伴生的鼠亚科及其动物群的层序对比[J]. 人类学学报, 2021, 40(06): 1041-1054. |
[8] | 王伟. 利用古蛋白技术分析巨猿演化地位的评述[J]. 人类学学报, 2020, 39(04): 717-726. |
[9] | 夏文婷;王社江;夏楠;鹿化煜;王先彦;孙雪峰;张红艳;张文超;卓海昕;邢路达;于青瑶;冯武明. 汉中盆地龙岗寺遗址第3地点出土的石制品[J]. 人类学学报, 2018, 37(04): 529-541. |
[10] | 刘连强;王法岗;杨石霞;岳健平. 泥河湾盆地马梁遗址第10地点2016年出土石制品研究[J]. 人类学学报, 2018, 37(03): 419-427. |
[11] | 裴树文;贾真秀;马东东;马宁;李潇丽. 泥河湾盆地麻地沟E5旧石器地点的遗址成因与石器技术[J]. 人类学学报, 2016, 35(04): 493-508. |
[12] | 王元;秦大公;严亚玲;金昌柱. 广西崇左泊岳山巨猿洞早更新世的鼠科化石[J]. 人类学学报, 2016, 35(04): 561-571. |
[13] | 王法岗. 泥河湾盆地南山边遗址发现的旧石器[J]. 人类学学报, 2016, 35(03): 331-342. |
[14] | 贾真秀;裴树文;马宁;张兴龙. 泥河湾盆地麻地沟E6和E7旧石器地点发掘简报[J]. 人类学学报, 2016, 35(03): 343-358. |
[15] | 邹松林;陈曦;张贝;赵克良;文军;邓里;同号文. 江西萍乡上栗县晚更新世哺乳动物化石发现[J]. 人类学学报, 2016, 35(01): 109-120. |
阅读次数 | ||||||
全文 |
|
|||||
摘要 |
|
|||||