人类学学报 ›› 2021, Vol. 40 ›› Issue (03): 469-489.doi: 10.16359/j.1000-3193/AAS.2021.0026
同号文1,2,3(), 张贝1,2,3, 陈曦4, 王晓敏5, 孙吉嘉1,2,3
收稿日期:
2020-10-27
修回日期:
2020-12-22
出版日期:
2021-06-15
发布日期:
2021-06-24
作者简介:
同号文,男,研究员,第四纪哺乳动物学专业。E-mail: 基金资助:
TONG Haowen1,2,3(), ZHANG Bei1,2,3, CHEN Xi4, WANG Xiaomin5, SUN Jijia1,2,3
Received:
2020-10-27
Revised:
2020-12-22
Online:
2021-06-15
Published:
2021-06-24
摘要:
狭义(或经典)泥河湾动物群主要产自桑干河北岸的下沙沟村周边,化石来源于不同地点;该动物群被作为我国北方早更新世的代表,但其绝对年代一直饱受争议,最新古地磁测年数据是2.2-1.7 MaBP。由于受河流及断层切割影响,泥河湾盆地内部不同区域尤其是桑干河两岸的地层对比,一直是个科学难题,其根本原因在于长期以来在桑干河南岸未发现重要化石点。最近,在桑干河南岸的岑家湾台地发现了山神庙咀化石点,目前已在此地点做过9次正规发掘,出土哺乳动物化石1526件,鉴定出25个种(含未定种),其中23种属于狭义泥河湾动物群的成员。由此看出,山神庙咀动物群的时代应当与狭义泥河湾动物群的时代接近,其地质年代应当不晚于1.7 MaBP。山神庙咀遗址与小长梁遗址隔沟相望,但其化石层稍高于小长梁文化层;山神庙咀动物群的研究结果还表明,小长梁遗址年代要比先前所测得的1.36 MaBP要早;小长梁周边的史前遗址的地质年代应当再做深入研究。小长梁地区的泥河湾层底部普遍存在砾石层,但其出现的水平位置以及砾石在分选和磨圆程度等方面都各不相同。调查清楚不同剖面中底砾层的分布状况及其性质,将会对本地区地层对比研究以及推断当时泥河湾盆地的古环境都将具有十分重要意义。
中图分类号:
同号文, 张贝, 陈曦, 王晓敏, 孙吉嘉. 泥河湾盆地早更新世山神庙咀遗址动物群及其时代意义[J]. 人类学学报, 2021, 40(03): 469-489.
TONG Haowen, ZHANG Bei, CHEN Xi, WANG Xiaomin, SUN Jijia. Chronological significance of the mammalian fauna from the Early Pleistocene Shanshenmiaozui site in Nihewan Basin, northern China[J]. Acta Anthropologica Sinica, 2021, 40(03): 469-489.
图2 山神庙咀化石点与周边遗址的位置关系 1.侏罗系顶面的夷平面,其上为更新统泥河湾层;2.山神庙咀旧石器遗址(1994年发掘区域);3.山神庙咀化石点;4.小长梁遗址A区;红色箭头指示滑坡处
Fig.2 Location of SSMZ and other prehistoric site in the vicinity
图3 山神庙咀化石点实测地层剖面图(A)及逐年发掘区域图(B)(A引自同号文等[2])
Fig.3 Profile of the strata at Shanshenmiaozui (A) and the grid system of the excavated area (B) (A is after Tong et al.[2])
图4 山神庙咀旧石器点与化石点的位置高差及探沟中出露的泥河湾层底界 A.山神庙咀旧石器遗址与化石点的位置关系;B.山神庙咀化石点发掘区域及探沟位置;C.探沟中暴露的地层序列;D.探沟底部暴露的底砾层;E.探沟中暴露的泥河湾层底部
Fig.4 Relative positions of the paleolithic site and fossil location at SSMZ as well as the exposed basal gravel in the trench
图5 小长梁地区各遗址底砾层出现位置(A-C)及特征(A’-C’) A’.小长梁遗址A区及底砾层;B-B’.山神庙咀化石点及底砾层;C-C’.照坡遗址及“石铺地面”(底砾层)
Fig.5 The feature of the basal gavels (A’-C’) and their horizons (A-C) at different sites in the XCL area
图7 山神庙咀化石点发现的小哺乳动物化石 1. Allactaga sibirica,右蹠骨(Mt II+III+IV) (V 26907);2-5. Ochotonoides complicidens,2. 右前颌骨带I1-2 (V 26905.1),3.右上颌骨带P2-M2 (V 26905.2),4.右下颌骨带p3-m3 (V 26905.3),5.左下颌骨带i1和p3-m3 (V 26905.4);6. Ochotona youngi,左下颌骨带i1和p3-m3 (V 26906)
Fig. 7 Micromammal fossils from SSMZ
图8 山神庙咀大型食草动物化石及与下沙沟有关化石比较 1a-1d.草原猛犸象,桡-尺骨(V 18010.89);2a-2c.巨副驼右侧蹠骨(V 26909.1);3a-3b.巨副驼左下颌骨(V 26909.2);4.秀丽日本鹿左角基部 (V 26910),前视;5a-5b.双叉麋鹿右侧掌骨(V 26911.2);6a-6b.双叉麋鹿右m3 (V 26911.1);7.秀丽黑鹿右角(A型);8.秀丽黑鹿右角(B型);9.秀丽黑鹿右角(C型);10.双叉麋鹿左下颌骨带p2-m3 (THP16. 875);11.秀丽黑鹿左下颌骨带p2-m3 (NIH 132);12.布氏真枝角鹿右下颌骨带p2-m3 (NIH 134)(水平翻转);13.布氏真枝角鹿左下颌骨带p2-m3 (V 25966.16)。7-8和10-11为第一作者在天津自然博物馆拍摄;12为第一作者在法国自然历史博物馆拍摄;9修改自Teilhard de Chardin and Piveteau.1930: Pl VIII-11[1]。1,2,4,5,6和13产自山神庙咀;3产自石沟;其它均产自下沙沟
Fig.8 Selected large herbivore fossils from SSMZ, compared with related fossils from Xiashagou (XSG)
狭义泥河湾动物群分类沿革Classic Nihewan Fauna | 山神庙咀动物群 SSMZ Fauna | 德马尼西动物群 Dmanisi Fauna | |||||
---|---|---|---|---|---|---|---|
Teilhard de Chardin[ | Barbour et al.[ | Teilhard de Chardin and Piveteau[ | 本文1)This paper | 本文This paper | Hemmer, et al.[ | ||
Erinaceus sp. | Erinaceus cf. dealbatus | Erinaceus cf. dealbatus | - | Beremendia fissidens | |||
Siphneus sp. | Siphneus tingi | Yangia tingi | - | - | |||
Arvicola sp. | Arvicolidae gen indet. | Borsodia chinensis | - | Mimomys tornensis | |||
Alactaga cf. annulata | Allactaga sibirica | Allactaga sibirica | |||||
Hystrix sp. | - | Hystrix sp. | |||||
Lagomys plicidens | Ochotona complicidens | Ochotonoides complicidens | Ochotonoides complicidens | - | |||
Ochotona youngi | Ochotona ex gr. Ochotona lagreli minor | ||||||
Canis megamastoides | Vulpes schlosseri | Canis (Nyctereutes) sinensis | Nyctereutes sinensis | Nyctereutes sp. | - | ||
Vulpes sp. | Vulpes sp. | Vulpes chikushanensis | - | Vulpes alopecoides | |||
Canis cf. lupus | Canis sp. | Canis chihliensis | Canis chihliensis | Canis chihliensis | Canis etruscus | ||
Canis chihliensis var. palmidens | Canis palmidens | - | - | ||||
Canis chihliensis var. minor | Eucyon minor | - | - | ||||
Ursus sp. | Ursus cf. etruscus | Ursus etruscus orientalis | - | Ursus etruscus | |||
Lutra sp. | Lutra licenti | Lutra licenti | - | - | |||
Mustela pachygnatha | Eirictis pachygnatha | - | - | ||||
Meles taxus | Meles taxus | Meles cf. leucurus | Meles chiai | - | Meles sp. | ||
Machairodus sp. | Machairodus sp. | Machairodus nihowanensis | Megantereon nihowanensis | ?Megantereon nihowanensis | Megantereon cultridens | ||
Homotherium crenatidens | - | Homotherium crenatidens | |||||
Cynailurus pleistocaenicus | Cynailurus pleistocaenicus | Acinonyx sp. | Acinonyx sp. | Acinonyx pardinensis | |||
Felis sp. | Felis sp. | Felis sp. | - | - | |||
Felis (Lynx) sp. | Lynx shansius | Lynx sp. | Lynx issiodorensis | ||||
?Panthera sp. | Panthera onca georgica | ||||||
Hyaena sp. | Chasmaporthetes progressus | - | - | ||||
Hyaena ultima | Hyaena ultima | Hyaena sinensis | Pachycrocuta licenti | Pachycrocuta licenti | Pachycrocuta perrieri Pachycrocuta sp. | ||
Crocuta honanensis | - | - | |||||
Elephas cf. trogontherii | Elephas namadicus | Mammuthus trogontherii | Mammuthus trogontherii | Mammuthus meridionalis | |||
Chalicotherid gen. indet. | Chalicotheridae gen. indet. | Postschizotherium chardini | - | - | |||
Hipparion sp. | Hipparion sinense | Hipparion (Proboscidipparion) sinense | Hipparion sp. | ||||
Equus sp. | Equus sp. | Equus sanmeniensis | Equus sanmeniensis | Equus sanmeniensis | Equus stenonis | ||
Equus teilhardi | - | - | |||||
Rhinoceros sp. | Rhinoceros cf. sinensis | Stephanorhinus yunchuchenensis | - | Stephanorhinus etruscus | |||
Rhinoceros cf. tichorhinus | Rhinoceros cf. tichorhinus | Coelodonta nihowansis | Coelodonta nihowansis | - | |||
Rhinocerid gen. indet. | Elasmotherium sp. | Elasmotherium sp. | Elasmotherium peii | - | |||
Circotherium sp. | Hesperotherium sp. | - | - | ||||
Sus sp. | Sus cf. lydekkeri | Sus lydekkeri | Sus lydekkeri | - | |||
Camelus (Paracamelus) gigas | Camelus (Paracamelus) gigas | Paracamelus gigas | Paracamelus gigas | - | |||
Cervus sp. | Cervulus cf. munjack | Cervulus cf. sinensis | Muntiacus bohlini | - | - | ||
Cervus cf. rusa | Cervus (Rusa) elegans | Nipponicervus elegans | Nipponicervus elegans | Dama (=Pseudodama =Cervus s.l.) cf. nestii | |||
Cervus cf. tetraceros | Cervus (Eucladoceros) boulei | Eucladoceros boulei | Eucladoceros boulei | Eucladoceros aff. tegulensis | |||
Cervus sp. | Cervus sp. | Cervus (Elaphurus) bifurcatus | Elaphurus bifurcatus | Elaphurus bifurcatus | - | ||
Antilope gen. indet. | Spirocerus wongi | Spirocerus wongi | Spirocerus wongi | Pontoceros sp. | |||
Spirocerus cf. peii | - | ||||||
?Antilospira robusta | - | ||||||
Gazella sp. | Gazella sp. | Gazella sinensis | Gazella sinensis | Gazella sinensis | - | ||
Gazella cf. subgutturosa | Gazella subgutturosa | - | - | ||||
Antilope gen. indet. | Hydropotes sp. | - | - | ||||
Ovis sp. | Ovis shantungensis | Ovis shantungensis | Ovis shantungensis | ||||
Ovis or Capra indet. | Ovis or Capra indet. | - | Capra dalii | ||||
Euceratherium (?) | Ovibovine gen. indet. | Megalovis piveteaui | Megalovis piveteaui | Soergelia cf. minor | |||
Leptobos sp. | Bison palaeosinensis | Bison palaeosinensis | Bison palaeosinensis | Bison (Eobison) georgicus |
表1 狭义泥河湾动物群组成及与山神庙咀动物群的对比
Tab.1 The taxonomic compositions of the Classic Nihewan Fauna and SSMZ Fauna
狭义泥河湾动物群分类沿革Classic Nihewan Fauna | 山神庙咀动物群 SSMZ Fauna | 德马尼西动物群 Dmanisi Fauna | |||||
---|---|---|---|---|---|---|---|
Teilhard de Chardin[ | Barbour et al.[ | Teilhard de Chardin and Piveteau[ | 本文1)This paper | 本文This paper | Hemmer, et al.[ | ||
Erinaceus sp. | Erinaceus cf. dealbatus | Erinaceus cf. dealbatus | - | Beremendia fissidens | |||
Siphneus sp. | Siphneus tingi | Yangia tingi | - | - | |||
Arvicola sp. | Arvicolidae gen indet. | Borsodia chinensis | - | Mimomys tornensis | |||
Alactaga cf. annulata | Allactaga sibirica | Allactaga sibirica | |||||
Hystrix sp. | - | Hystrix sp. | |||||
Lagomys plicidens | Ochotona complicidens | Ochotonoides complicidens | Ochotonoides complicidens | - | |||
Ochotona youngi | Ochotona ex gr. Ochotona lagreli minor | ||||||
Canis megamastoides | Vulpes schlosseri | Canis (Nyctereutes) sinensis | Nyctereutes sinensis | Nyctereutes sp. | - | ||
Vulpes sp. | Vulpes sp. | Vulpes chikushanensis | - | Vulpes alopecoides | |||
Canis cf. lupus | Canis sp. | Canis chihliensis | Canis chihliensis | Canis chihliensis | Canis etruscus | ||
Canis chihliensis var. palmidens | Canis palmidens | - | - | ||||
Canis chihliensis var. minor | Eucyon minor | - | - | ||||
Ursus sp. | Ursus cf. etruscus | Ursus etruscus orientalis | - | Ursus etruscus | |||
Lutra sp. | Lutra licenti | Lutra licenti | - | - | |||
Mustela pachygnatha | Eirictis pachygnatha | - | - | ||||
Meles taxus | Meles taxus | Meles cf. leucurus | Meles chiai | - | Meles sp. | ||
Machairodus sp. | Machairodus sp. | Machairodus nihowanensis | Megantereon nihowanensis | ?Megantereon nihowanensis | Megantereon cultridens | ||
Homotherium crenatidens | - | Homotherium crenatidens | |||||
Cynailurus pleistocaenicus | Cynailurus pleistocaenicus | Acinonyx sp. | Acinonyx sp. | Acinonyx pardinensis | |||
Felis sp. | Felis sp. | Felis sp. | - | - | |||
Felis (Lynx) sp. | Lynx shansius | Lynx sp. | Lynx issiodorensis | ||||
?Panthera sp. | Panthera onca georgica | ||||||
Hyaena sp. | Chasmaporthetes progressus | - | - | ||||
Hyaena ultima | Hyaena ultima | Hyaena sinensis | Pachycrocuta licenti | Pachycrocuta licenti | Pachycrocuta perrieri Pachycrocuta sp. | ||
Crocuta honanensis | - | - | |||||
Elephas cf. trogontherii | Elephas namadicus | Mammuthus trogontherii | Mammuthus trogontherii | Mammuthus meridionalis | |||
Chalicotherid gen. indet. | Chalicotheridae gen. indet. | Postschizotherium chardini | - | - | |||
Hipparion sp. | Hipparion sinense | Hipparion (Proboscidipparion) sinense | Hipparion sp. | ||||
Equus sp. | Equus sp. | Equus sanmeniensis | Equus sanmeniensis | Equus sanmeniensis | Equus stenonis | ||
Equus teilhardi | - | - | |||||
Rhinoceros sp. | Rhinoceros cf. sinensis | Stephanorhinus yunchuchenensis | - | Stephanorhinus etruscus | |||
Rhinoceros cf. tichorhinus | Rhinoceros cf. tichorhinus | Coelodonta nihowansis | Coelodonta nihowansis | - | |||
Rhinocerid gen. indet. | Elasmotherium sp. | Elasmotherium sp. | Elasmotherium peii | - | |||
Circotherium sp. | Hesperotherium sp. | - | - | ||||
Sus sp. | Sus cf. lydekkeri | Sus lydekkeri | Sus lydekkeri | - | |||
Camelus (Paracamelus) gigas | Camelus (Paracamelus) gigas | Paracamelus gigas | Paracamelus gigas | - | |||
Cervus sp. | Cervulus cf. munjack | Cervulus cf. sinensis | Muntiacus bohlini | - | - | ||
Cervus cf. rusa | Cervus (Rusa) elegans | Nipponicervus elegans | Nipponicervus elegans | Dama (=Pseudodama =Cervus s.l.) cf. nestii | |||
Cervus cf. tetraceros | Cervus (Eucladoceros) boulei | Eucladoceros boulei | Eucladoceros boulei | Eucladoceros aff. tegulensis | |||
Cervus sp. | Cervus sp. | Cervus (Elaphurus) bifurcatus | Elaphurus bifurcatus | Elaphurus bifurcatus | - | ||
Antilope gen. indet. | Spirocerus wongi | Spirocerus wongi | Spirocerus wongi | Pontoceros sp. | |||
Spirocerus cf. peii | - | ||||||
?Antilospira robusta | - | ||||||
Gazella sp. | Gazella sp. | Gazella sinensis | Gazella sinensis | Gazella sinensis | - | ||
Gazella cf. subgutturosa | Gazella subgutturosa | - | - | ||||
Antilope gen. indet. | Hydropotes sp. | - | - | ||||
Ovis sp. | Ovis shantungensis | Ovis shantungensis | Ovis shantungensis | ||||
Ovis or Capra indet. | Ovis or Capra indet. | - | Capra dalii | ||||
Euceratherium (?) | Ovibovine gen. indet. | Megalovis piveteaui | Megalovis piveteaui | Soergelia cf. minor | |||
Leptobos sp. | Bison palaeosinensis | Bison palaeosinensis | Bison palaeosinensis | Bison (Eobison) georgicus |
图9 狭义泥河湾动物群组成及其地质分布 蓝色标记的属种也出现在山神庙咀化石点;橙色标记的地点有古人类化石发现;绿色区域代表狭义泥河湾动物群和山神庙咀动物群主要分布时段;各属种的分布时限主要参考邱占祥[29],Qiu[52]及Rook and Martínez-Navarro[56]
Fig.9 The up-dated taxonomic compositions of the Classic Nihewan Fauna
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